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Quackgrass Biology Outline & Bibliography


Prepared by: Bruce Battles, April 1998

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Quackgrass (Elytrigia repens)


Prepared by: Bruce Battles


 Introduction

A. Name

 Family: POACAE- Grass Family

 Genus: Elytrigia

 Species: repens

 Common Names: quackgrass and couch grass

 Previously named: Agroypyron repens, Triticum repens and Elymus repens

B. General

 perennial

 reproduce by seeds and rhizomes

 native of Europe

 introduced to New England during colonization

 probably introduced to Quebec during same period

 rapid spreading westward with European settlement

 spread through infested bromegrass seed and infested hay and straw

C. Geographical Distribution

 Europe, Australia, New Zealand

 temperate zones of Asia and North and South America

 found in all states of the United States of America

 found from Newfoundland to British Columbia in Canada (Boivin 1967)

 Taxonomy (Plant Morphology and Anotomy)

A. Identification

 Blade

 Sheath

 Ligule

 Culm

 spikes

 Spikelets

 Awns

 Glumes

 Rhizomes

B. Morphology

 growth composed of aerial shoots and rhizomes

 horizontal rhizome growth beneath soil surface in spring and summer

 axillary buds at base of aerial shoots renew rhizome growth

 axillary buds along rhizomes are mainly dormant

 aerial shoots form at end of growing season from erect (vertical) rhizome tip

 adventitious roots form at nodes of rhizome

C. Phenology

 seeds germinate early spring

 tiller production (2-3 months after emergence)

 rhizome production--------June to August

 vertical rhizome growth and shoot development occur in Autumn

 mature plant will form the following year

 flowering--------------------June to July

 seed ripening---------------August to September

 seed drop-------------------late September

 natural seasonal cycle may be altered by cultural practices and environment

  Reproduction

A. Floral Biology

 cross pollinator

 wind-pollinated

 virtually self-sterile

 great variation found from seed reproduction

 many biotypes produced from cross pollination

B. Seed Production and dispersal

 seed production range: 15-400 seeds per plant stem (commonly 25-40)

 variability in seed production dependent on: (Williams and Attwood 1970, 1971)

a.) variability will be greater at edge of large stands produced by one clone (Palmer and Sagar 1963)

b.) closer exposure to other genotype will produce greatest variability

 seeds fall passively from parent plants

C. Viability of seeds and germination

 alternating temperatures are required for germination (Chepil 1946; Andersen 1968; Palmer and Sagar 1963)

 no after ripening period

 dormancy of 2-3 years (Carder 1961)

 viability: maximum of 4 years (Brackney and Seely 1966)

 germination promoted by gibberellic acid (Holm and Miller 1972)

 germination inhibited by abscissic acid (Holm and Miller 1972)

D. Vegetative reproduction and dispersal

 apical dominance in rhizomes

 parent plant cloning

(Raleigh et al. (1962) reported 3 meter diameter spread from parent plant)

( Raleigh et al. (1962) reported up to 200 shoots)

 increased reproduction vegetatively as result of soil disturbance (tillage)

 can be dispersed through tillage (lodging of rhizomes in implements)

 longevity of rhizome buds: 2 years (Lemieux, et al. 1993)

 buds can exhibit dormancy

 rhizome bud viability increases with increasing distance from the primary shoot (Dekker and Chandler 1985)

  Habitat

A. Climatic requirements

 cool season grass

 Common in areas of moderate rainfall (rare in low rainfall areas)

B. Substratum

 wide variety of soil types

 pH range: 4.5-8.0 (Doyon 1965, 1968; Rousseau 1968)

 high salt tolerance: 70-95 mg/100g of soil (Tesu et al. 1972)

 has been found on saline soils

 rhizome growth is reduced in compacted soils (Wolcott and Carlson 1947)

C. Communities which quackgrass occurs

 mainly occuring in areas where native community has been disturbed

 common in:

 competition

  Population Dynamics

 first growing season: extensive subtillering (clump growth habit)

 as many as 150 rhizomes or rhizome branches formed first year

 second growing season: clump growth pattern develops into patches and later continuous stands

 plants produced from tillers are more vigorous and aggressive than plants from seeds

 biomass from nonliving quackgrass is believed to be allelopathic

 high growth rates during cool periods

 fierce competitor for nutrients: percentage of total nutrients removed from soil by July and total kg/ha removed by 2,000 kg shoot dry weight per hectare respectively: (Bandeen and Buchholtz 1967)

 Economic Importance

A. Detrimental

 competes strongly with cultivated crops

 reduces yields: (up to)

 increased ear moisture of corn at harvest

 delayed tasseling and silking of corn (Bandeen and Bucholtz 1967)

 highly competitive for nutrients

 reduced value of grass seed crops from contamination

 host for several grain crop pests

 

B. Beneficial

 preferred plant for rangeland

 can be used for pasture and hay

 total crude protein comparable to forages such as Timothy grass

 rhizomes are efficient soil binders for erosion prevention

 provides cover for wildlife animals

 dried and ground rhizomes can serve as a source of flour

 has many useful properties for use in herbals

  Control Strategies

 minimum of 2 years total control for eradication (2 year rhizome viability) (Lemiux et al. 1991)

 prevention:

 repeated tillage:

 herbicidal control: (corn)

 herbicidal control: (soybeans)

  Significance of Quackgrass as a problem weed

 spreads quickly by rhizomes

 difficult to control with normal tillage (usually will distribute it further)

 fierce competitor for nutrients

 limited herbicides for use in certain crops

 use as a forage has increased the dispersal (although will serve as a fair forage)

 


Quackgrass Bibliography

 Table of Contents

 Biology and Habitat

 Seed Production

 Dormancy

 Growth and Development

 Competition

 Economic Importance

 Management and Control (other than herbicides)

 Herbicidal Control

 Variability (hybrids and biotypes)

 Biology and Habitat

Arny, A. C. 1932. Varieations in the organic reserves in underground parts of five perennial weeds from late April to November. Minn. Agric. Exp. Sta. Tech. Bull. 84: 1-28.

Andersson,-A Is it couchgrass? [Agropyron repens] 1973

Andersson,-A Couchgrass, an unnecessary weed problem. [Agropyron repens] 1972

Conn,-J.S. Seed viability and dormancy of 17 weed species after burial for 4.7 years in Alaska. Weed-Sci. Champaign, Ill. : Weed Science Society of America. Mar 1990. v. 38 (2) p. 134-138.

Conn,-J.S.; Deck,-R.E. Seed viability and dormancy of 17 weed species after 9.7 years of burial in Alaska. Weed-sci. Champaign, Ill. : Weed Science Society of America. Oct/Dec 1995. v. 43 (4) p. 583-585.

Friebe,-A.; Schulz,-M.; Kuck,-P.; Schnabl,-H. Phytotoxins from shoot extracts and root exudates of Agropyron repens seedlings. Phytochemistry-Oxford. Oxford : Elsevier Science Ltd. Mar 1995. v. 38 (5) p. 1157-1159.

Gillespie,-G.R.; Vitolo,-D.B. Response of quackgrass (Elytrigia repens) biotypes to primisulfuron. Weed-technol. Champaign, Ill. : The Weed Science Society of America. Apr/June 1993. v. 7 (2) p. 411-416.

Greub,-L.-J; Collins,-M; Carlson,-S.-K Relationship of morphological characteristics to forage quality in quackgrass Crop-Science. v. 26 July/Aug. '86 p. 819-22

Hagin,-R.D.; Bobnick,-S.J. Isolation and identification of a slug-specific molluscicide from quackgrass (Agropyron repens, L. Beauv.). 1991

Hakansson,-Sigurd Quackgrass, biology and control. Uppsala : Aktuellt fran lantbrukshogskolan, 1977. 31 p. : ill. –

Hunter,-J.H..; Hsiao,-A.I.; McIntyre,-G.I Effect of nitrogen on the glyphosate-induced inhibition of rhizome bud growth in quackgrass (Elytrigia repens). Weed-sci. Champaign, Ill. : Weed Science Society of America. July/Sept 1993. v. 41 (3) p. 426-433.

Korhammer,-S.A.; Haslinger,-E. Isolation of a biologically active substance from rhizomes of quackgrass [Elymus repens (L.) Gould]. J-agric-food-chem. Washington, D.C. : American Chemical Society. Sept 1994. v. 42 (9) p. 2048-2050.

Lemieux,-C.; Cloutier,-D.C.; Leroux,-G.D. Distribution and survival of quackgrass (Elytrigia repens) rhizome buds. Weed-science (USA). (Oct-Dec 1993). v. 41(4) p. 600-606.

Lemieux,-C.; Cloutier,-D.C.; Leroux,-G.D. Sampling quackgrass (Elytrigia repens) populations. Weed-Sci. Champaign, Ill. : Weed Science Society of America. Oct/Dec 1992. v. 40 (4) p. 534-541.

Muller,-K.; Karch,-K. Couchgrass biology studies for effective control Economic crops, Agropyron repens. Untersuchungen zur Biologie der Quecke als Grundlage einer effektiven Bekampfung. 1981

Reidy,-M.E.; Swanton,-C.J. Response of four quackgrass (Elytrigia repens (L.) Nevski) biotypes to desiccation. Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. July 1994. v. 74 (3) p. 643-646.

Robertson, J. M. 1989. Apical dominance in rhizomes of quackgrass: inhibitory effect of scale leaves. Weed Sci 37: 680-687.

Smith,-B.S.; Pawlak,-J.A.; Murray,-D.S.; Verhalen,-L.M.; Green,-J.D. Interference from established stands of silverleaf nightshade (Solanum elaeagnifolium) on cotton (Gossypium hirsutum) lint yield. Weed-Sci. Champaign, Ill. : Weed Science Society of America. Mar 1990. v. 38 (2) p. 129-133.

Stobbe,-E.H. Biology of quackgrass [Agropyron repens]. Proceedings-of-the-Annual-Meeting-North-Central-Weed-Control-Conference (USA). (1976). v. 31 p. 151-152.

Swanson,-M.B.; Davis,-G.A.; Kincaid,-L.E.; Schultz,-T.W.; Bartmess,-J.E.; Jones,-S.L.; George,-E.L. A screening method for ranking and scoring chemicals by potential human health and environmental impacts. Environ-toxicol-chem. Pensacola, Fla. : SETAC Press. Feb 1997. v. 16 (2) p. 372-383.

Tardif,-F.J.; Leroux,-G.D. Rhizome bud viability of quackgrass (Elytrigia repens) treated with glyphosate and quizalofop. Weed-Technol-J-Weed-Sci-Soc-Am. Champaign, Ill. : The Society. July/Sept 1990. v. 4 (3) p. 529-533.

Taylor,-J.S.; Robertson,-J.M.; Harker,-K.N.; Bhalla,-M.K.; Daly,-E.J.; Pearce,-D.W.

Apical dominance in rhizomes of quackgrass, Elytrigia repens: the effect of auxin, cytokinins, and abscisic acid. Can-j-bot. Ottawa : National Research Council of Canada, 1951-. Feb 1995. v. 73 (2) p. 307-314.

Werner, P. A., R. Rioux. 1977. The biology of Canadian weeds. 24. Agropyrom repens (L.) Beauv. Can J Plant Sci. 57: 905-919.

Zollinger,-R.K.; Penner,-D.; Kells,-J.J. Absorption, translocation, and foliar activity of clopyralid and tribenuron in perennial sowthistle (Sonchus arvensis). Weed-Sci. Champaign, Ill. : Weed Science Society of America. Oct/Dec 1992. v. 40 (4) p. 528-533.

 Seed Production

Williams, E.D. and Attwood, P.J. 1970 Seed production by couch grass (Agropyron repens). N.A.A.S.Q. Rev. 89:42-46.

Williams, E.D. and Attwood, P.J. 1971. Seed production of Agropyron repens (L.) Beauv. In arable crops in England and Wales in 1969. Weed Res. 11:22-30.

 Dormancy

Conn,-J.S.; Deck,-R.E. Seed viability and dormancy of 17 weed species after 9.7 years of burial in Alaska. Weed-sci. Champaign, Ill. : Weed Science Society of America. Oct/Dec 1995. v. 43 (4) p. 583-585.

Conn,-J.S. Seed viability and dormancy of 17 weed species after burial for 4.7 years in Alaska. Weed-Sci. Champaign, Ill. : Weed Science Society of America. Mar 1990. v. 38 (2) p. 134-138.

Dekker, J. H., Chandler. 1985. Herbicide effect on the viability of quackgrass rhizome buds. Can J Plant Sci 65: 1057-1064.

Smith,-B.S.; Pawlak,-J.A.; Murray,-D.S.; Verhalen,-L.M.; Green,-J.D. Interference from established stands of silverleaf nightshade (Solanum elaeagnifolium) on cotton (Gossypium hirsutum) lint yield. Weed-Sci. Champaign, Ill. : Weed Science Society of America. Mar 1990. v. 38 (2) p. 129-133.

Swanson,-M.B.; Davis,-G.A.; Kincaid,-L.E.; Schultz,-T.W.; Bartmess,-J.E.; Jones,-S.L.; George,-E.L. A screening method for ranking and scoring chemicals by potential human health and environmental impacts. Environ-toxicol-chem. Pensacola, Fla. : SETAC Press. Feb 1997. v. 16 (2) p. 372-383.

 Growth and Development

Chancellor, R.J. 1974. The development of dominance amongst shoots arising from fragments of Aggropyron repens rhizomes. Weed Res. 14:29-38.

McIntyre, G. I. 1965. Some effects of the nitrogen supply on the growth and development of Agropyron repens (L.)Beauv. Weed. Res. 5:1-12

McIntyre, G. I. 1969. Apical dominance in the rhizome of Agropyron repens. Evidence of competition for carbohydrate as a factor in th mechanism of inhibition. Can. J. Bot. 47:1189-1197.

McIntyre, G. I. 1987. Studies on the growth and development of Agropyron repens: interaction effects of humidity, calcium, and nitrogen on growth of the rhizome apex and lateral buds. Can J Bot 6€: 1427-1432

McIntyre, G.I. 1970. Studies on bud development in the rhizome of Agropyron repens. The influence of temperature, light intensity and bud position on the pattern of development. Can. J. Bot. 48:1903-1909.

Palmer, J.H. 1963. Agropyron repens (L) Beauv. The seasonal development and growth of the parent plant and rhizome. New Phytol. 57:145-159.

Rogan, P.G. and Smith, D.L. 1975a. The effet of temperature and nitrogen level on the morphology of Agropyron repens (L.) Beauv. Weed Res. 15:93-99.

Rogan, P.G. and Smith, D.L. 1975b. Rates of leaf initiation and leaf growth in Agropyron repens (L.) Beauv. J. Exp. Bot. 26:70-78.

Rogan, P.G. and Smith, D.L. 1976. Experimental control of bud inhibition in rhizomes of Agropyron repens (l.) Beauv. Z. Pflanzenphysiol. 78:113-121.

Williams, E.D. 1970a. Effects of decreasing th light intensity on the growth of Agropyron repens L. Beauve. In the field. Weed Res. 10:360-366.

Williams, E.D. 1970b. Studies on the growth of seedlings of Agropyron repens (L.) Veauv. And Agrostis gigantea Roth. Weed Res. 10:321-330.

Williams, E.D. 1971. Effects of light intensity photoperiod and nitrogen on the growth of seedlings of Agropyron repens L. Beauv. And Agrostis gigantea Roth. Weed Res. 11:159-170.

Williams, E.D. 1973a. A comparison of the growth and competition behaviour of seedlings and plants from rhizomes of Agropyron repens L. Beauv. And Agrostis gigantea Roth. Weed Res. 13:422-429.

Williams, E.D. 1975. Growth of seedlings of Agropyron repens L. Beauv. And Agrostis gigantea Roth in cereal crops. Weed Res. 15:299-306.

 Competition

Bandeen, J.D. and Buchholtz, K.P. 1967. Competitive effects of quack grass upon corn as modified by fertilization. Weeds 15:220-224

Friesen, G. And Shebeski, L. H. 1960. Economic losses caused by weed competition in Manitoba grain fields. 1. Weed species, their relative abundance and their effect on crop yield. Can. J. Plant Sci 40:457-467.

Pageau,-D.; Tremblay,-G.F. Effect of quackgrass on the productivity of barley seeded at different seeding rates and row spacings. Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. July 1995. v. 75 (3) p. 613-618.

Sikkema, P. H., J. Dekker. 1987. Use of infrared thermometry in determining critical stress periods induced by quackgrass (Agropyron repens in soybeans). Weed Sci 35: 784-791.

Taylor, D. R., L. W. Aarssen. 1988. An interpretation of phenotypic plasticity in Agropyrom repens (Graminae). Am J Bot 75: 401-413.

Welbank, P.J. 1961. A study of the nitrogen and water factors in competition with Agropyron repens (L.) Beauv. Ann. Bot. 25:116-137.

Weston, L. A., A. R. Putnam. 1985. Inhibition of growth, nodulation, and nitrogen fixation of legumes by quackgrass. Crop Sci 25: 561-565.

Weston, L. A., 1986. Inhibition of legume seedling growth by residues and extracts of quackgrass (Agropyron repens). Weed Sci 34: 366-372.

Williams, E.D. 1973a. A comparison of the growth and competition behaviour of seedlings and plants from rhizomes of Agropyron repens L. Beauv. And Agrostis gigantea Roth. Weed Res. 13:422-429.

Young, F. L., D. L. Wyse, and R. J. Jones. 1984. Quackgrass (Agropyrom repens) interference on corn (Zea Mays). Weed Sci 32: 226-234.

 Economic Importance

Asay,-K.H.; Dewey,-D.R. Improvement of range grasses through interspecific hybridization with quackgrass (Elytrigia repens L. Nevski). Repr-U-S-Dep-Agric-Agric-Res-Serv. Washington, D.C. : The Service. 1985. [519] 2 p.

Friesen, G. And Shebeski, L. H. 1960. Economic losses caused by weed competition in Manitoba grain fields. 1. Weed species, their relative abundance and their effect on crop yield. Can. J. Plant Sci 40:457-467

Hoffman,-P.C.; Sievert,-S.J.; Shaver,-R.D.; Welch,-D.A.; Combs,-D.K. In situ dry matter, protein, and fiber degradation of perennial forages. J-dairy-sci. Champaign, Ill. : American Dairy Science Association. Sept 1993. v. 76 (9) p. 2632-2643.

Narasimalu,-P.; McRae,-K.B.; Kunelius,-H.T. The composition and utilization of silage prepared from timothy (Phleum pratense L.) and quackgrass (Elytrigia repens (L.) Neoski). 1989

Neuteboom,-J.H.; Elzebroek,-A.T.G. Productivity and nutritive value of couchgrass Elytrigia repens andEnglish ryegrass Lolium perenne with different mowing frequencies Digestibility. Produktiviteit en voederwaarde van kweek en Engels raaigras bij verschillende maaifrequenties. 1980

Seoane,-J.R.; Christen,-A.M.; Veira,-D.M.; Fontecilla,-J. Performance of growing steers fed quackgrass hay supplemented with canola meal. 1992

Sheaffer,-C.C.; Wyse,-D.L.; Marten,-G.C.; Westra,-P.H. The potential of quackgrass for forage production. J-prod-agric. [Madison, WI] : American Society of Agronomy, c1987-. Apr/June 1990. v. 3 (2) p. 256-259.

 

 Management and Control (other than herbicides)

Albrecht,-K.A.; Djajanegara,-A.S. Use of cultural practices to manage quackgrass in alfalfa without herbicides. Proceedings Progress in Wisconsin sustainable agriculture March 1990 /. [Wisconsin?] : University of Wisconsin, [1990?]. p. 124-126.

Brimhall, P. B. 1970. Modifications in the weed flora due to weed control techniques. Proc. 2nd Int. Mett. Selective Weed Control Beet Crops. Rottedam. 249 pp.

Canada Department of Agriculture. 1944. Control of couch grass in Manitoba. Can. Dep. Agric. Publ. 765.

Carder, A. C. 1961. Couchgrass control in Alberta. Alta. Dep. Agric. Publ. 149.

Derksen,-D.A.; Thomas,-A.G.; Lafond,-G.P.; Loeppky,-H.A.; Swanton,-C.J.

Impact of agronomic practices on weed communities: fallow within tillage systems.

Weed-sci. Champaign, Ill. : Weed Science Society of America. Apr/June 1994. v. 42 (2) p. 184-194.

Doll,-J.D. Quackgrass management in field crops. North-Cent-reg-ext-publ. East Lansing, Mich. : Cooperative Extension Service. Aug 1993. (219) 4 p.

Hahn,-R.R. Quackgrass control programs change for field corn. 1991

Loeppky,-H.A.; Derksen,-D.A. Quackgrass suppression through crop rotation in conservation tillage systems. Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. Jan 1994. v. 74 (1) p. 193-197.

Moyer,-J.R.; Schaalje,-G.B. Quackgrass (Elytrigia repens) interferance and control in seed alfalfa (Medicago sativa). Weed-technol. Champaign, Ill. : The Weed Science Society of America. Jan/Mar 1993. v. 7 (1) p. 58-64.

Pageau,-D. Effect of barley seeding rate and row spacing on control of quackgrass.Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. Oct 1996. v. 76 (4) p. 921-926.

Salzman,-F.; Renner,-K.; Kells,-J. Controlling quackgrass. Ext-Bull-E-Coop-Ext-Serv-Mich-State-Univ. East Lansing, Mich. : The Service. Dec 1992. (2251,rev.) 2 p.

 Herbicidal Control

Akhavien, A. A. 1971. Effects of 2-chloroethylphosphonic acid (ethrel) and selected environmental factors on growth of quack grass (Agropyron repens L (Beauv.)) and field bindweed (Convolvulus arvensis L.). Ph.D. Thesis. Oregon State University.

Baird, D.D. and Begeman, G. F. 1972. Post emergence characterization of a new quack grass herbicide. Proc. North East. Weed Sci. Soc. 26: 100-106.

Brackney, C. T. And Seely, C. I. 1966. Quack grass control in Idaho. Agric. Exp. Sta. Bull. 455.

Bhowmik,-P.C.; O'Toole,-B.M.; Andaloro,-J. Effects of nicosulfuron on quackgrass (Elytrigia repens) control in corn (Zea mays). Weed-Technol-J-Weed-Sci-Soc-Am. Champaign, Ill. : The Society. Jan/Mar 1992. v. 6 (1) p. 52-56.

Chandler,-K.; Murphy,-S.D.; Swanton,-C.J. Effect of tillage and glyphosate on control of quackgrass (Elytrigia repens). Weed-technol. Champaign, Ill. : The Weed Science Society of America. July/Sept 1994. v. 8 (3) p. 450-456.

Curran,-W.S.; Werner,-E.L.; Hartwig,-N.L. Effectiveness of herbicides and tillage on quackgrass (Elytrigia repens) control in corn (Zea mays). Weed-technol. Champaign, Ill. : The Weed Science Society of America. Apr/June 1994. v. 8 (2) p. 324-330.

Darwent,-A.L.; Clayton,-G.W.; Drabble,-J.C.; Mills,-P.F.; Wolynetz,-M.S. Integration of glyphosate and quizalofop with tillage for quackgrass (Elytrigia repens) management in continuous annual crop and legume plowdown rotations. Weed-technol. Champaign, Ill. : The Weed Science Society of America. Oct/Dec 1996. v. 10 (4) p. 923-930.

Dekker, J. H., Chandler. 1985. Herbicide effect on the viability of quackgrass rhizome buds. Can J Plant Sci 65: 1057-1064.

Gaynor,-J.D.; Hamill,-A.S. Timing of atrazine application for control of quackgrass (Agropyron repens). Phytoprotection. Quebec : Quebec Society for the Protection of Plants, 1963-. 1993. v. 74 (2) p. 89-99.

Gillespie,-G.R. Basis for the differential response of quackgrass (Elytrigia repens) biotypes to primisulfuron. Weed-sci. Champaign, Ill. : Weed Science Society of America. Jan/Mar 1994. v. 42 (1) p. 8-12.

Gillespie,-G.R.; Vitolo,-D.B. Response of quackgrass (Elytrigia repens) biotypes to primisulfuron. Weed-technol. Champaign, Ill. : The Weed Science Society of America. Apr/June 1993. v. 7 (2) p. 411-416.

Gronwald,-J.W.; Jourdan,-S.W.; Wyse,-D.L.; Somers,-D.A.; Magnusson,-M.U.

Effect of ammonium sulfate on absorption of imazethapyr by quackgrass (Elytrigia repens) and maize (Zea mays) cell suspension cultures. Weed-sci. Champaign, Ill. : Weed Science Society of America. July/Sept 1993. v. 41 (3) p. 325-334.

Hamill,-A.S.; Zhang,-J. Quackgrass control with glyphosate and SC-0224 in corn and soybean. Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. Jan 1995. v. 75 (1) p. 239-299.

Harker, K. N., J. Dekker. 1988. Effects of phenology on translocation patterns of several herbicides in quackgrass. Weed €c; 36: 463-47€

Harker,-K.N. Short-term split application effects of grass-specific herbicides on quackgrass (Elytrigia repens) under field conditions. Weed-technol. Lawrence, Kans. : The Weed Science Society of America. Oct/Dec 1995. v. 9 (4) p. 710-715.

Harker,-K.N.; O'Sullivan,-P.A. Herbicide comparisons on quackgrass (Elytrigia repens) within different crop competition and tillage conditions. Weed-sci. Champaign, Ill. : Weed Science Society of America. Jan/Mar 1993. v. 41 (1) p. 94-99.

Harker,-K.N.; Taylor,-J.S. Chlormequat chloride (CCC) pretreatments may enhance quackgrass (Elytrigia repens) control with sethoxydim. Weed-technol. Champaign, Ill. : The Weed Science Society of America. July/Sept 1994. v. 8 (3) p. 499-507.

Hunter,-J.H.; Hsiao,-A.I.; McIntyre,-G.I. Effect of nitrogen on the glyphosate-induced inhibition of rhizome bud growth in quackgrass (Elytrigia repens). Weed-sci. Champaign, Ill. : Weed Science Society of America. July/Sept 1993. v. 41 (3) p. 426-433.

Ivany,-J.A. Effect of haloxyfop on quackgrass (Elytrigia repens) and potatoes (Solanum tuberosum). Weed-Technol-J-Weed-Sci-Soc-Am. Champaign, Ill. : The Society. Jan/Mar 1991. v. 5 (1) p. 72-75.

Leroux,-G.D. Effect of initial quackgrass density on control with glyphosate prior to spring wheat. Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. Oct 1993. v. 73 (4) p. 1283-1291.

Leroux,-G.D.; Hamel,-G. Influence of ethylan and ammonium sulfate on glyphosate phytotoxicity to quackgrass (Elytrigia repens). Adjuvants for agrichemicals / edited by Chester L Foy. Boco Raton, FL : CRC Press, c1992.. p. 127-136.

Maruska,-D.W.; Lym,-R.G.; Messersmith,-C.G. Quackgrass control in cropland with various spring-applied herbicides. Res-Prog-Rep-West-Soc-Weed-Sci. [S.l.] : The Society. 1993. p. III/96-III/97.

McMullan,-P.M. Influence of temperature after spraying on glyphosate control of quackgrass (Elytrigia repens (L.) Nevski) previously exposed to light frost. Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. July 1994. v. 74 (3) p. 667-669.

Reidy,-M.E.; Swanton,-C.J. Postemergence control of quackgrass [Elytrigia repens (L.) Nevski] with DPX-79406 in corn (Zea mays L.). Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. Apr 1994. v. 74 (2) p. 375-381.

Ruiter,-H.-de; Meinen,-E. Adjuvant-increased glyphosate uptake by protoplasts isolated from quackgrass Elytrigia repens (L.) Nevski. Weed-sci. Champaign, Ill. : Weed Science Society of America. Jan/Mar 1996. v. 44 (1) p. 38-45.

Ruiter,-H.-de; Uffing,-A.J.M.; Meinen,-E. Influence of surfactants and ammonium sulfate on glyphosate phytotoxicity to quackgrass (Elytrigia repens). Weed-technol. Champaign, Ill. : The Weed Science Society of America. Oct/Dec 1996. v. 10 (4) p. 803-808.

Shieh,-W.J.; Geiger,-D.R.; Buczynski,-S.R. Distribution of imported glyphosate in quackgrass (Elytrigia repens) rhizomes in relation to assimilate accumulation. Weed-sci. Champaign, Ill. : Weed Science Society of America. Jan/Mar 1993. v. 41 (1) p. 7-11.

Tardif,-F.J.; Leroux,-G.D. Translocation of glyphosate, quizalofop, and sucrose in quackgrass (Elytrigia repens) biotypes. Weed-sci. Champaign, Ill. : Weed Science Society of America. July/Sept 1993. v. 41 (3) p. 341-346.

Twidwell,-E.K.; Kephart,-K.D.; Clay,-S.A. Quackgrass control in established alfalfa with sethoxydim. Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. July 1994. v. 74 (3) p. 647-651.

Wallace,-R.W.; Bellinder,-R.R. Rust-infected quackgrass (Elytrigia repens) growth and rhizome bud death after glyphosate application. Weed-sci. Champaign, Ill. : Weed Science Society of America. July/Sept 1993. v. 41 (3) p. 501-507.

 Variability (hybrids and biotypes)

Asay,-K.H.; Dewey,-D.R. Improvement of range grasses through interspecific hybridization with quackgrass (Elytrigia repens L. Nevski). Repr-U-S-Dep-Agric-Agric-Res-Serv. Washington, D.C. : The Service. 1985. [519] 2 p.

Buchholtz, K.P. 1958. Variations in the sensitivity of clones of quackgrass to dalapon. Proc. North Cent. Weed Control Conf. 15:18-19.

Dewey, D.R. 1965. Synthetic hybrids of new world and old world agropyrons. 11. Agropyron repens X Agropyron riparium. Amer. J. Bot. 52:1039-1045

Konarev,-A.V. Occurrence of genetic material from Elytrigia juncea (L.) Nevski (2n = 14) in polyploid quackgrass species. 1981

Neuteboom, J. H. 1975. Variability of Elytrigia repens L. Beauv. On Dutch agricultural soils. Meded. Landbouwhogesch. Wageningen 7:1-29

Neuteboom, J.H. 1981. Effect of different mowing regimes on the growth and development of four clones of couch (Elytrigia repens L. Desv. ; syn Agropyron repens L. Beauv.) in monocultures and in mixtures with perennial ryegrass (Lolium perenne L.) . Meded. Landbouwhogesch. Wageningen 15:1-26.

Rahman,-M.S. Relationship between visual colour rating and chlorophyll content, photosynthetic rate and some growth characteristics in couchgrass (Cynodon spp. L.) Varieties. 1983

Reidy,-M.E.; Swanton,-C.J. Response of four quackgrass (Elytrigia repens (L.) Nevski) biotypes to desiccation. Can-j-plant-sci. Ottawa : Agricultural Institute of Canada, 1957-. July 1994. v. 74 (3) p. 643-646.

Tardif, F.J.; Leroux, G.D. 1991. Variability of quackgrass (Agropyron repens) biotypes in Quebec. Phytoprotection 72:115-121.

Tardif,-F.J.; Leroux,-G.D. Response of three quackgrass biotypes to nitrogen fertilization. 1992

Tardif,-F.J.; Leroux,-G.D. Variability of quackgrass (Agropyron repens) biotypes in Quebec. Phytoprotection. Saint-Hyacinthe : Quebec Society for the Protection of Plants. 1991. v. 72 (3) p. 115-121.

Westra, P. H.; Wyse, D.L. 1981. Growth and Development of Quackgrass (Agropyron repens) Biotypes. Weed Science29:44-52.

Williams, E.D. 1973, Variations in growth of seedlings and clones of Agropyron repens (L) Beauv. Weed Res. 13:24-41.


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